Definitely the lifetime warranty will come into effect. Only time will tell. Hopefully no real serious issues will arise once the razor is in my hands. Plus, I've never been afraid of a little experimentation. Good, clean shaves is what I'm looking forward to, with a Feather or the optional Gem blade. Let's see how it goes.....The rails don't hold the blade at all, the locking nubs do. So any wearing of the rails will not affect how the razor holds the blade.
I have an early V1, and when I first bought it, the rails were rather tight on blade insertion/extraction. After some use, these rails "broke in", and changing the blade was easier, but still locked in the blade just as securely as it always has. The danger with using a GEM in V1 that is not clipped at it's sides is that it does not lock in, pushes against, and can possibly wear away the locking nubs. To prevent against that possibility, clip your GEM's. If you don't clip the GEM's in V1, you take that risk upon yourself in not using the razor as it was designed.
With V2, the blade doesn't slide against the rails at all. The locks hold the blade more loosely, yet still securely. There shouldn't be any wear from friction here, because there really isn't any, and even if there were, the V2 OneBlade head is made from PANACEA stainless steel, more than twice as hard and durable than any other stainless steel, including the steel used in blades.
In any case, OneBlade's "forever" warranty will cover any manufacturing defect, or natural wear and tear issues. I don't think you'd need this warranty anyway, but I'm not sure they'd cover damaged locking nubs from using an un-clipped GEM in V1.